These creatures are similar to Unmarked Spectralids but discovering a fray of these is much more unpleasant. They swoop down and swarm your Pikmin squad, eating unprotected Pikmin. They are just as frail as Unmarked Spectralids, so if you throw like a madman you'll probably take out the cloud of unwanted moths. They are most effective against you when you're fighting other enemies and they can be a huge pain.
In Pikmin: Return
||Fenestari prismatus nocturnista
Twilight Spectralids appear in Pikmin: Return, although they behave much differently. They still fly around and produce the same noises as other spectralids, but they will glow pink when in a cave or outside near sunset. Twilight Spectralids take two hits from Pikmin to defeat, and when they are hit, they aggressively squeal and flap their wings harder to shower attackers below them in poisonous powder, which can poison and kill all Pikmin except White Pikmin. Upon death, a cloud of poison dust will burst from the Twilight Spectralid as its body falls from the ground. It can be brought back to the Onion for five seeds if it is given to the Winged Pikmin, and it will give three seeds to any other color. A larger, more dangerous variant of this enemy is known as the Greater Twilight Spectralid.
- "This spectralid has evolved a natural defense against predators by making its scales poisonous. These scales can also be conveniently scattered around the air if it is being swarmed by predators. These scales, unfortunately, are harmful if inhaled by most other lifeforms, including pikmin, hocotations, and koppaites. These spectralids can also take two direct hits by pikmin while airborne. Another interesting thing about these spectralids is that they let off a dim, purplish glow that attracts winged pikmin."
- "Inedible. Will cause severe stomach aches... ouch..."
- "This remarkable spectralid gives off a dim pink glow. It is quite beautiful when in a dim environment. Speaking of environments, this spectralid chooses a dark cave or a hollow tree trunk to reside in, in contrast to most other spectralids, which choose bright areas to live in or on."
- "This spectralid must be contained in a very secure and airtight container, as its poisonous powder might contaminate any cargo that is near it... It also flaps around violently when captured, as if trying to escape or attack... maybe this is its form of attacking..."